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In addition, although equilibrium concentrations of inorganic carbon species are established by the pH, catalysis to equilibrium by CA ensures that diffusion of CO sub across membranes, and diffusion of all inorganic carbon species within the aqueous cellular environment, is not limited by flux rates between species. Proposed photosynthetic roles for the highly abundant stromal enzyme betaCA have included facilitation of CO sub movement across the chloroplast envelope and maintenance of maximal rates of CO sub and HCO sub sup diffusion through the stroma by rapidly equilibrating C sub i speciation Badger and Price.